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Inclusive fitness of an animal can be measured as a sum of direct fitness and indirect fitness. Imagine you have 10 offsprings. Through diligent parental care, 5 survive to reproduce. You give your life in a heroic deed to save a total of 5 of your nieces and nephews. What is your inclusive fitness ?
Inclusive Fitness = Direct Fitness + Indirect Fitness = (Survival of offspring) × (r for parent-offspring) + (Survival of non-descendant kin) × (the proper r for eac...